You are an excellent assistant at answering questions based on Spanish texts. You will be given an article and a question, and you must answer the question based only on information in the article. The response should be extracted from the context verbatim.

After thoroughly reading the article and analyzing the question, respond with the answer.

Title:
Force

Context:
All of the forces in the universe are based on four fundamental interactions. The strong and weak forces are nuclear forces that act only at very short distances, and are responsible for the interactions between subatomic particles, including nucleons and compound nuclei. The electromagnetic force acts between electric charges, and the gravitational force acts between masses. All other forces in nature derive from these four fundamental interactions. For example, friction is a manifestation of the electromagnetic force acting between the atoms of two surfaces, and the Pauli exclusion principle, which does not permit atoms to pass through each other. Similarly, the forces in springs, modeled by Hooke's law, are the result of electromagnetic forces and the Exclusion Principle acting together to return an object to its equilibrium position. Centrifugal forces are acceleration forces that arise simply from the acceleration of rotating frames of reference.:12-11:359

Question:
What do gravitational forces act between?

masses